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Sea buckthorn (Hippophae rhamnoides L.) is an important perennial shrub, a pseudo-fruit crop in China. The plants have a high nutritional value because of its high content of vitamin C and polyphenols. In recent years, sea buckthorn has been promoted for its high economic value and is quickly growing in China. However, powdery mildew caused by golovinomyces cichorii has been observed in Xinjiang and Hebei provinces since 2008. The disease occurs mainly in the autumn and winter and causes severe yield losses, especially in the case of highly infested cultivars. Five cultivars of sea buckthorn, ‘Feu’, ‘Henan’, ‘Hainan’, ‘Hainan 1’, and ‘Hainan 3’, were used to study the symptom development and the progression of disease. A typical infection stage in plants showed that powdery mildew covered the entire leaf surfaces, resulting in stunted growth. As the disease progressed, the leaflets of infected leaves separated from the leaf veins and turned discolored and necrotic. Eventually, the leaves turned brown and collapsed. Microscopic observation of infected leaves revealed typical powdery mildew infection structures. Conidiophores were filiform, 0.5 to 1.3 × 5 to 20 μm, bearing four to 20 conidia per cell. Conidiophores were initially erect, but became more laterally flexuous or decumbent after maturity. Phialides were somewhat flexuous, measuring 11 to 20 μm. Conidia were ellipsoid, 2 to 4 × 2 to 3 μm, with an average of 2.3 μm. The pathogen was identified by morphological characteristics (1,2,3,4). A total of 20 isolates were deposited in the China General Microbiological Culture Collection Center (No.CGMCC3.19584). For mating experiments, two strains, M-177 and M-201, were inoculated on the

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